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While coach Rasi Erasmus has named a more conventional set of replacements, with five specialist forwards instead of the seven he has tested in the past, former Harlequins center Andre Osterhuizen has been deployed as a flanker off the bench in the past.
Cheslin Kolbe and Damien Willems will both win their 50th caps as part of the starting line-up.
“England are a quality team, and we expect them to give everything they have in this match, so it was important to select a squad with experience combined with some young players who have proven they have what it takes to perform at this level,” said Erasmus, whose side beat a Barbarian side 80-31 in a warm-up match on June 2.
The England squad is expected to be announced later this week.
South Africa: Williams; Kolbe, Kriel, de Allende, Arendse; Libock, Williams; Nche, Marx, T du Toit, Etzebeth, Nortje, Kolisi, PS du Toit, Wiese
Replacement: Wessels, Steenkamp, Porten, Van Staden, Hannekom, Reinch, Esterhuizen, Moody
